Fish were hand-fed four times daily (08:00 a.m, 11:00 p.m, 14:00 p.m and 17:00 p.m respectively) with different diets at a rate of 10% of their body weight (BW) during the first 28 days of feeding and 5% for the rest of the feeding period, with feeding ration being adjusted in accordance to weight gain of fish after random sampling at 14 days